Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Appeltans W., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D., eds (2011). Species 2000 & ITIS Catalogue of Life, 5th December 2011. Digital resource at www.catalogueoflife.org/col/. Species 2000: Reading, UK.